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PASADENA, Calif. • Sean Mannion passed for a career-high 379 yards and two touchdowns, Oregon State put the clamps on No. 19 UCLA's offense and the Beavers opened their Pacific 12 Conference season by topping the Bruins 27-20 on Saturday.

The victory was the 74th at Oregon State for coach Mike Riley, matching the school record set by Lon Stiner, the Beavers' coach from 1933 to 1948. Riley is in his 12th year with Oregon State.

Mannion, a third-year sophomore, completed 24 of 35 passes with one interception and has passed for at least 200 yards in 13 straight games.

Oregon State's Markus Wheaton had nine receptions for 150 yards and has caught at least one pass in 25 consecutive games. Brandin Cooks had six catches for 175 yards, and Storm Woods rushed for 96 yards on 21 carries for the Beavers.

The Bruins (3-1, 0-1 Pac-12) scored 15 touchdowns and averaged 622.0 yards to rank second nationally in total offense in their previous three games. But the Beavers (2-0, 1-0) held them to 444 yards and two TDs.

Oregon State had played just once in the season's first three weeks, beating then-No. 13 Wisconsin 10-7 on Sept. 8. The season opener against Nicholls State was postponed because of concerns over Hurricane Isaac.

UCLA's Johnathan Franklin, the country's leading rusher with a 180.3-yard average, was held to 45 yards on 12 carries. He averaged 8.2 yards per carry this year entering the game. Redshirt freshman Brett Hundley completed 27 of 42 passes for 372 yards and a touchdown. —